The fragmentation of tropical forests in eastern Brazil as a result of agricultural expansion and other human activities has decimated biodiversity even within the pockets of forest that still remain, a new study has found. Using wildlife surveys and interviews conducted at 196 forest fragments across a 253,000-square-kilometer region inside Brazil’s Atlantic Forest, a team of researchers estimated that only about 22 percent of the animals that once inhabited the region are still there — far lower than earlier estimates. According to their findings, published in the journal PLoS ONE, white-lipped peccaries have been “completely wiped out,” while jaguars, lowland tapirs, woolly spider-monkeys and giant anteaters are essentially extinct. The loss of wildlife has even extended to areas where forest canopies are still relatively intact, said Carlos Peres, an ecologist at the University of East Anglia and lead author of the study. While the Atlantic Forest once covered more than 1.5 million square kilometers, about 90 percent has been cleared for agriculture, pasture, or urban expansion. Most remaining patches of forest, researchers say, are about the size of a football field.